Officer of the Order of Canada

• Awarded on: April 22, 1993
• Invested on: February 16, 1994

Professor of medical biochemistry at the University of Calgary, he is at the forefront of biomedical research. His early research into the regeneration of insulin activity has contributed to our understanding of protein structure and function. However, he is best known for his ground-breaking research into the process of spermatogenesis in trout, which has revolutionized our understanding of chromosome structure.